Color Modification
Chocolate (be)
Alternative Names: Brown, Liver

Chocolate color discovered in the Lancashire Heeler, also known as "be," is a variant found in the TYRP1 gene, known as the B locus. This variant is associated with a dog’s darkest color pigment being limited to brown instead of black. Meaning all dark hair, nails and skin (including eye rims, nose and paw pads) will be a shade of brown or chocolate. Eye color is also affected, and is typically gold or amber.
More Info
Did you know?
Chocolate coloration is often termed "liver" in Lancashire Heelers. And, more specifically, the recognized color pattern is called "liver and tan" for the breed.
How it works
To show chocolate coloration a dog must inherit two chocolate variants, one from each parent. This can either be two copies of a particular variant, such as this one ("be”), or two of any combination of chocolate variants.
Technical Details
Gene | TYRP1 |
---|---|
Also Called | Brown (B) Locus |
Variant | T>G |
Chromosome | 11 |
Coordinate | 33,326,719 |
All coordinates reference CanFam3.1
References & Credit
Credit to our scientific colleagues:
Wright, H.E, Schofield, E., Mellersh, C.S, Burmeister, L.M. (2019). A novel TYRP1 variant is associated with liver and tan coat colour in Lancashire Heelers. Anim Genet 50(6), 783. View the article